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		<H1>Publications</H1>
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		<H2>Peer-reviewed:</H2>[# = graduate student; % = undergraduate student]
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Gladstone, N.S. #, Pieper, E.B., <B>Keenan, S.W.</B>, Paterson, A.T., Slay, M., Dooley, K., Engel, A.S., and Niemiller, M.L. (2021). Discovery of the Blue Ridge Springsnail, <I>Fontigens orolibas</I>, Hubricht, 1957 (Gastropoda: Emmericiidae) in east Tennessee and its conservation implications. <I>Freshwater Mollusk Biology and Conservation</I>,<U> in press</U>.

<LI>

Emmons, A.L.#, Mundroff, A.Z., <B>Keenan, S.W.</B>, Davoren, J., Andronowski, J., Carter, D. O., and DeBruyn, J.M. (2020). Characterizing the postmortem human bone microbiome from surface decomposed remains. <I>PLoS One</I>, 15:e0218636, DOI:10.1371/journal.pone.0218636.
<LI>
Fanter, C.E.#, Lin, Z., <B>Keenan, S.W.</B>, Janzen, F.J., Mitchell, T.S., and Warren, D.E. (2020). Development-specific transcriptomic profiling suggests new mechanisms for anoxic survival in the ventricle of overwintering turtles. <I>Journal of Experimental Biology</I>, 223, DOI: 10.1242/jeb.213918.
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<B>Keenan, S.W.</B>, Schaeffer, S. M., and DeBruyn, J.M. (2019). Spatial changes in soil stable isotopic composition in response to carrion decomposition. <I>Biogeosciences</I>, 16:3929-3939, DOI: 10.5194/bg-16-3929-2019.
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<B>Keenan, S.W.</B> and DeBruyn, J. M. (2019). Changes to vertebrate tissue stable isotope (&delta;<sup>15</sup>N) composition during decomposition. <I>Scientific Reports</I>, 9:9929, DOI: 10.1038/s41598-019-46368-5.
<LI>
<B>Keenan, S.W.</B>, Pasteris, J.D., Wang, A., and Warren, D.E. (2019). Heterogeneous
bioapatite carbonation in western painted turtles is unchanged after anoxia. <I>Comparative
Biochemistry and Physiology, Part A</I>, 233: 74-83. 
<LI>
<B>Keenan, S.W.</B>, Emmons, A.L.#, Taylor, L.S.#, Phillips, G., Mason, A.R.#, Mundorff, A., Bernard, E.C., Davoren, J., and DeBruyn, J.M. (2018). Spatial impacts of a multi-individual grave on microbial and microfaunal communities and soil biogeochemistry. <I>PLoS One</I>, DOI: 10.1371/journal.pone/0208845.
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Odegard, D.T.#, Sonnenfelt, M.A.%, Bledsoe, J.G., <B>Keenan, S.W.</B>, Hill, C.A.#, and Warren, D.E. (2018). Changes in the material properties of the shell during simulated aquatic hibernation in the anoxia-tolerant painted turtle. <I>Journal of Experimental Biology</I>, DOI: 10.1242/jeb.176990.  <BR></LI>
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<B>Keenan, S.W.</B>, Schaeffer, S.M., Jin, V.L., and DeBruyn, J.M. (2018). Mortality hotspots: nitrogen cycling in forest soils during vertebrate decomposition. <I>Soil Biology and Biochemistry</I>, 121: 165-176. <BR></LI>
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<B>Keenan, S.W.</B>, Widga, C., DeBruyn, J.M., and Schaeffer, S.M. (2018). Nutrient hotspots through time: a field guide to modern and fossil taphonomy in East Tennessee in Engel, A.S. and Hatcher, B., eds., <I>Field Guide 50: 2018 Southeastern Section Meeting Field Guide</I>, pp. 61-74. <BR></LI>
<LI>

<B>Keenan, S.W.</B> and Engel, A.S. (2017). Reconstructing diagenetic conditions of bone at the Gray Fossil Site, Tennessee, USA. <I>Palaeogeography, Palaeoclimatology, Palaeoecology</I>, 471: 48-57.  <BR></LI>
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<B>Keenan, S.W.</B> and Engel, A.S. (2017). Early diagenesis and recrystallization of bone. <I>Geochimica et Cosmochimica Acta</I>, 196: 209-223. <BR></LI>
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<B>Keenan, S.W.</B> and Tellez, M. (2016). Trachemys scripta elegans (Red-eared slider) and Chrysemys picta bellii (Western painted turtle). Endoparasites. <I>Herpetological Review</I>, 47(3): 451-453. <BR></LI>
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<B>Keenan, S.W.</B> (2016). From bone to fossil: a review of the diagenesis of bioapatite. <I>American Mineralogist</I>, 101: 1943-1951. <BR></LI>
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<B>Keenan, S.W.</B>, Engel, A.S., Roy, A., and Bovenkamp-Langlois, G.L. (2015). Evaluating the consequences of diagenesis and fossilization on bioapatite lattice structure and composition. <I>Chemical Geology</I>, 413: 18-27. <BR></LI>
<LI>

<B>Keenan, S.W.</B>, Hill, C.A., Kandoth, C., Buck, L.T., and Warren, D.E. (2015). Transcriptomic response of the western painted turtle heart and telencephalon to anoxia. <I>PLoS One</I>, DOI: 10.1371/journal.pone.0131669. <BR></LI>
<LI>

<B>Keenan, S.W.</B> and Elsey, R.M. (2015). The good, the bad, and the unknown: microbial symbioses of the American alligator. <I>Integrative and Comparative Biology</I>, DOI: 10.1093/icb/icv006. <BR></LI>
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<B>Keenan, S.W.</B>, Niemiller, M.L., and Williams, B.W. (2014). Observations of Cambarus bartonii cavatus (Decapoda: Cambaridae) and ectosymbiotic branchiobdellidans (Annelida: Clitellata) in Cruze Cave, Knox County, Tennessee, USA. <I>Speleobiology Notes</I>, 6: 55-61. <BR></LI>
<LI>

<B>Keenan, S.W.</B> and Ellwood, B.B. (2014). Geophysical evaluation of the Richland and Holloway Mounds, southeastern Louisiana, U.S.A. <I>Geoarchaeology</I>, 29: 312-325.   <BR></LI>
<LI>

<B>Keenan, S.W.</B> and Scannella, J.B. (2014). Paleobiological implications of a Triceratops bonebed from the Hell Creek Formation, Garfield County, northeastern Montana. In (Wilson, G.P., Clemens, W.A., Horner, J.R., and Hartman, J.H., eds.) <I>Through the end of the Cretaceous in the type locality of the Hell Creek Formation in Montana and adjacent areas</I>. Geological Society of America Special Paper 503. Pp.349-364. <BR></LI>
<LI>

<B>Keenan, S.W.</B>, Engel, A.S. and Elsey, R.M. (2013). The alligator gut microbiome and implications for archosaur symbioses. <I>Scientific Reports</I>, DOI: 10.1038/srep02877. <BR></LI>

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		<H2>
		Other Publications:
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<a href=" "><SMALL></SMALL></A><B>Keenan, S.W.</B> (2019). Fossilization. <I> Encyclopedia of Geology, 2nd Edition</I>, Elsevier (in press).
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<B>Keenan, S.W.</B>, Paterson, A.T., Niemiller, M.L., Slay, M.E., Clark, S.A., and Engel, A.S. (2017). Observations of the first stygobiont snail (Hydrobiidae, Fontigens sp.) in Tennessee. <I>Proceedings of the 17th International Congress of Speleology</I>, Sydney, Australia, pp. 91-94. <BR></LI>
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Engel, A.S., Niemiller, M.L., Zigler, K.S., Stephen, C.D.R, Carter, E.T., Paterson, A.T., <B>Keenan, S.W.</B>, and Taylor, S.J. (2017). Invertebrate and vertebrate cave fauna records for the Appalachian Valley and Ridge. <I>Proceedings of the 17th International Congress of Speleology</I>, Sydney, Australia, pp. 82-86.
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<B>Keenan, S.W.</B> (2013). Freshwater vertebrate animal metagenomics, Alligatorinae. In (Nelson, K., ed.) <I>Encyclopaedia of Metagenomics</I>. Springer, DOI: 10.1007/978-1-4614-6418-1_101-8.
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		<H2>
		Abstracts (last five years):
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<LI>
DeVault, G.A.#* and <B>Keenan, S.W.</B> (2020). Using tooth wear and decomposition to interpret geochemistry in paleontological localities. <I> Geological Society of America</I> (online, poster, *presenter).
<LI>
<B>Keenan, S.W.</B> and DeBruyn, J. M. (2020). Surface and subsurface vertebrate decomposition result in protracted soil nitrogen stable isotopic enrichment. <I>Rocky Mountain Section of Geological Society of America</I>, Provo, UT (May, oral presentation, cancelled due to COVID).
<LI>
Cyrier, M.%* and <B>Keenan, S.W.</B> (2020). Impacts of seasonality on subsurface conditions in Rushmore Cave, South Dakota. <I>Rocky Mountain Geobiology</I>, Golden, CO (*presenter, cancelled due to COVID).
<LI>
<B>Keenan, S.W.</B> and DeBruyn, J. M. (2019). Quantifying the long-term impacts of animal decomposition on soil biogeochemical cycling. <I>Geological Society of America</I>, Phoenix, AZ (poster)
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<B>Keenan, S.W.</B> (2019). Temporal and spatial dynamics of animal decomposition in terrestrial systems. <I>Geobiology Symposium</I>, Banff, Canada (<U>invited oral presentation</U>).
<LI>

Driebergen, J.#* and <B> Keenan, S.W. </B> (2019). Cosmopolitan anuran assemblages: proxies for the effects of climate change? <I>Society of Vertebrate Paleontology</I>, Brisbane, Australia (*presenter).
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Emmons, A.L.#*, <B>Keenan, S.W.</B>, Davoren, J., DeBruyn, J.M., Carter, D., and Mundorff, A.Z. (2019). DNA preservation in bone in a multi-individual burial. <I>Proceedings of the American Academy of Forensic Sciences</I>, Baltimore, MD (*presenter). <BR></LI>
<LI>
Mason, A.R.#*, <B>Keenan, S.W.</B>, Emmons, A.L.#, Taylor, L.S.#, Phillips, G., Mundorff, A.Z., Bernard, E.C., and DeBruyn, J.M. (2019). Spatial impact of a multi-individual grave on soil biochemistry and microbial ecology. <I>Soil Science Society of America</I>, San Diego, CA (*presenter). <BR></LI>
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<B>Keenan, S.W.</B> and DeBruyn, J.M. (2018). Spatial and temporal soil responses to carrion decay. <I>Goldschmidt</I>, Boston, MA (poster).
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Emmons, A.L.#*, <B>Keenan, S.W.</B>, Mason, A.#, Holand, K., Campagna, S., Davoren, J., DeBruyn, J.M, Mundorff, A. (2018). Edaphic and biogenic influences of skeletal DNA degradation in a multi-individual grave. <I>International Society for Microbial Ecology</I>, Leipzig, Germany (*presenter) <BR></LI>
<LI>

Mason, A.R.#*, <B>Keenan, S.W.</B>, Emmons, A.L.#, Taylor, L.S.#, Phillips, G., Mundorff, A.Z., Bernard, E.C., and DeBruyn, J.M. (2018). Spatial impact of a multi-individual grave on soil biochemistry and microbial ecology. <I>American Society for Microbiology</I>, Atlanta, GA (*presenter; <U>winner of the ASM outstanding abstract award</U>) <BR></LI>
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Grimes, E.%*, <B>Keenan, S.W.</B>, and DeBruyn, J.M. (2018). The enzymatic and geochemical composition of bone. <I>Undergraduate Research Symposium and EUReKA (Exhibition of Undergraduate Research and Creative Achievements)</I>, University of Tennessee (*oral and poster presentations). <BR></LI>
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<B>Keenan, S.W.</B> and DeBruyn, J.M. (2018). Temperature and microbial effects on nitrogen cycling at simulated mortality decomposition hotspots. <I>Southeastern Geological Society of America</I>, Knoxville, TN (oral presentation) <BR></LI>
<LI>

<B>Keenan, S.W.</B>, Widga, C., and DeBruyn, J.M. (2018). Reconstructing nutrient hotspots in time: insights from the Gray Fossil Site, Tennessee. <I>Southeastern Geological Society of America</I>, Knoxville, TN (oral presentation).  <BR></LI>
<LI>

Emmons, A.L.#*, <B>Keenan, S.W.</B>, Taylor, L.S.#, Davoren, J., DeBruyn, J.M., Phillips, G., Bernard, E.C., and Mundorff, A.Z. (2018). Microbial ecology and soil geochemistry in a multi-individual grave. <I>Proceedings of the American Academy of Forensic Sciences</I>, Seattle, WA (*presenter).
 <BR></LI>
<LI>

<B>Keenan, S.W.</B>, Schaeffer, S.M, Jin, V.L., and DeBruyn, J.M. (2017). Soil biogeochemical cycling and isotope dynamics during vertebrate decomposition. <I>Soil Science Society of America</I>, Tampa, FL (oral presentation). <BR></LI>
<LI>

Taylor, L.S.#*, Phillips, G., <B>Keenan, S.W.</B>, Bernard, E.C., and DeBruyn, J.M. (2017). Nematode community succession in vertebrate carcass decomposition soil hot spots. <I>Soil Science Society of America</I>, Tampa, FL (*presenter). <BR></LI>
<LI>

<B>Keenan, S.W.</B>, Schaeffer, S.M., and DeBruyn, J.M. (2017). Stable isotope dynamics of vertebrate tissues during decomposition. <I>International Society for Environmental Biogeochemistry</I>, Cairns, Australia (oral presentation). <BR></LI>
<LI>

Bernard, E.C.*, Phillips, G., Taylor, L.S.#, <B>Keenan, S.W.</B>, and DeBruyn, J.M. (2017). Nematode assemblages and succession on and under beaver carcasses. <I>Society of Nematologists</I>, Williamsburg, VA (*presenter).
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<LI>

Taylor, L.S.#*, Phillips, G., <B>Keenan, S.W.</B>, Bernard, E.C., and DeBruyn, J.M. (2017). Spatial nematode community distributions associated with human decomposition in a mass grave. <I>Society of Nematologists</I>, Williamsburg, VA (*presenter).
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<LI>

Engel, A.S.*, Niemiller, M.L., Zigler, K.S., Stephen, C.D.R, Carter, E.T., Paterson, A.T., <B>Keenan, S.W.</B>, and Taylor, S.J. (2017). Invertebrate and vertebrate cave fauna records for the Appalachian Valley and Ridge. <I>Proceedings of the 17th International Congress of Speleology</I>, Sydney, Australia, pp. 82-86 (*presenter). <BR></LI>
<LI>

<B>Keenan, S.W.</B>, Paterson, A.T., Niemiller, M.L., Slay, M.E., Clark, S.A., and Engel, A.S.* (2017). Observations of the first stygobiont snail (Hydrobiidae, Fontigens sp.) in Tennessee. <I>Proceedings of the 17th International Congress of Speleology</I>, Sydney, Australia, pp. 91-94 (*presenter).
<BR></LI>
<LI>

<B>Keenan, S.W.</B>, Schaeffer, S.M., Jin, V.L., and DeBruyn, J.M. (2017). Biogeochemistry of vertebrate decomposition in a forest ecosystem. <I>Southeastern Biogeochemistry Symposium</I>, Athens, GA. <BR></LI>
<LI>

Emmons, A.L.#*, <B>Keenan, S.W.</B>, Mundorff, A., Davoren, J., and DeBruyn, J.M. (2017). Skeletal DNA preservation and bone-associated microbes: the implications for DNA sampling strategies in forensic identification. <I>Proceedings of the American Academy of Forensic Sciences</I>, New Orleans, LA, p. 108-109 (*presenter).
 <BR></LI>
<LI>

DeBruyn, J.M.*, Cobaugh, K.L., Moats, M., Stevens, J.D.#, and <B>Keenan, S.W.</B> (2016). Postmortem hot spots: soil microbial community succession below decomposing vertebrate mortalities. <I>Soil Science Society of America</I>, Phoenix, AZ (*presenter).
<BR></LI>
<LI>

<B>Keenan, S.W.</B>, Hauther, K.A.#, and DeBruyn, J.M. (2016). Soil responses to a nutrient &quot;hotspot&quot;: the biogeochemistry of vertebrate decomposition in a forest ecosystem. <I>Midwest Geobiology</I>, Cincinnati, OH (oral presentation). <BR></LI>
<LI>

<B>Keenan, S.W.</B>, Emmons, A.L.#, Mundorff, A., Davoren, J.M., and DeBruyn, J.M. (2016). Bone-associated microbes: implications for the long-term stability of bone in terrestrial systems. <I>Geological Society of America</I>, Denver, CO (oral presentation). <BR></LI>
<LI>

Engel, A.S.*, Paterson, A.T., and <B>Keenan, S.W.</B> (2016). A little drop will do: surprises from epikarstic drip water microbiology. <I>National Speleological Society Convention</I>, Ely, NV (*presenter). <BR></LI>
<LI>

<B>Keenan, S.W.</B>, Paterson, A.T., Niemiller, M.L., Slay, M.E., and Engel, A.S. (2016). The first record of a stygobiontic cave snail (Hydrobiidae) from Tennessee. <I>National Speleological Society Convention</I>, Ely, NV (oral presentation). <BR></LI>
<LI>

<B>Keenan, S.W.</B>, and Engel, A.S. (2015). Thermodynamic predictions of bioapatite stability in environmental systems. <I>GSA Abstracts with Programs</I>, 47(7):461, session no. 180, Baltimore, MD (oral presentation). <BR></LI>
<LI>

<B>Keenan, S.W.</B>, Pasteris, J.D., Wang, A., and Warren, D.E. (2015). A mineralogical perspective on a physiological problem: bioapatite carbonation in the western painted turtle. <I>Midwest Geobiology Symposium</I>, Bloomington, IN (oral presentation).
<BR></LI>
<LI>

Engel, A.S.*, Harmon, D.%, Paterson, A.T., and <B>Keenan, S.W.</B> (2015). Cave dripwater microbiology reveals epikarstic biogeochemical cycling that can affect speleothem records. <I>International Society of Environmental Biogeochemistry</I>, Slovenia (*presenter).<BR></LI>
<LI>

<B>Keenan, S.W.</B>+, Pasteris, J.D., Wang, A., and Warren, D.E. (2015). Mineralogical contributions to overwintering survival in the western painted turtle. <I>St. Louis Ecology, Evolution, and Conservation</I>, St. Louis, MO (+invited oral presentation). <BR></LI>
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<IMAGE SRC='myimages/bone geochem.jpg' WIDTH=180 BORDER="0" ALIGN=center TITLE=“Bones provide a wealth of info, including geochemsitry!“ >
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<IMAGE SRC='myimages/skeleton in field.jpg' WIDTH=180 BORDER="0" ALIGN=center TITLE=“Beaver skeleton weathering in the field, East Tennessee, US.“  >
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<IMAGE SRC='myimages/DSCN5943.jpg' WIDTH=180 BORDER="0" ALIGN=center TITLE=“Hell Creek Formation, Montana.“  >
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<IMAGE SRC='myimages/snail.jpg' WIDTH=180 BORDER="0" ALIGN=center TITLE=“Stygobiont snail, East Tennessee, US.“  >	
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